I've had a really really close look under bright light and subdued light and I think you are right - it's definitely a grain direction issue rather than an adhesion issue. From the top or across the grain, however close you look, there is no discernible issue. In fact it shimmers like a great tight piece of proper vertically grained lacquered spruce does.
However, if you look obliquely from either end along the grain, the areas of difference are as plain as the stripes on a freshly lawn-mowered and rolled lawn.
Clearly, I have sanded differently or to a different level or finished in a different direction in those areas somewhere in the grain levelling process. Whether the egg white has caused or exacerbated the issue, I can't tell.
The back and sides look OK, so I will continue with those as normal. (the Lacewood is a very smooth tight grained, almost formica-like, wood so no grain issues there). The top will be sanded down to the wood, maybe this evening, and re-done.